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This situation caused a renewed interest in the Ornithocheirus material and the validity of the several names based on it, for it might be possible that it could by more detailed studies be established that the Brazilian pterosaurs were actually junior synonyms of the European types.
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Based on these specimens, Cope named two new species, Ornithochirus umbrosus and Ornithochirus harpyia, in an attempt to assign them to the large European genus Ornithocheirus, though he misspelled the name (forgetting the 'e').
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This would distinguish both from the species Criorhynchus simus and justify a revival of the genus Coloborhynchus that since an analysis by Reginald Walter Hooley in 1914 had generally been considered identical to the genus Criorhynchus or the genus the latter had again been sunk into, Ornithocheirus.
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Referring to Seeley's designation of 1881 he considered Ornithocheirus simus, holotype CAMSM B.54428, to be the type species.
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"Ornithocheirus" buenzeli (often mis-spelled O. bunzeli) is a pterosaur species known from parts of a humerus (upper arm bone) and part of a lower jaw found in late Cretaceous period (Campanian) rocks of the Grünbach Formation, Austria.
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However, in 2013 Taissa Rodrigues and Alexander Wilhelm Armin Kellner concluded firstly that Pterodactylus fittoni Owen 1859 was not a part of the genus Anhanguera and secondly that Ornithocheirus nasutus was not identical to it regardless.
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Owen was not an evolutionist and he therefore considered the name Ornithocheirus to be inappropriate; he also thought it was possible to distinguish two main types within the material, based on differences in snout form and tooth position — the best fossils consisted of jaw fragments.
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Furthermore, Kuhn was the first to formally designate C. clavirostris as the type species of the genus, rather than one of the Ornithocheirus species.
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Criorhynchus, "ram beak", consisted entirely of former Ornithocheirus species: the type species Criorhynchus simus and furthermore C. eurygnathus, C. capito, C. platystomus, C. crassidens and C. reedi.
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As a result, though over forty species have been named in the genus Ornithocheirus over the years, only O. simus is currently considered valid by all pterosaur researchers.
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It has at times been synonymised with Coloborhynchus, Criorhynchus, Lonchodectes, or Ornithocheirus.
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In 1888 Edward Newton renamed several existing species names into Ornithocheirus, as Ornithocheirus clavirostris, O. daviesii, O. sagittirostris, O. validus, O. giganteus, O. clifti, O. diomedeus, O. nobilis, O. curtus, O. umbrosus, O. harpyia, O. macrorhinus and O. hlavaci.
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However, Hooley's classification was rarely applied later in the century, when it became common to subsume all the poorly preserved and confusing material under the name Ornithocheirus.
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Later the species was sometimes assigned to other genera, resulting in the new combinations Ornithocheirus validus (Newton 1888) and Cycnorhamphus validus (Owen 1870).
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Tropeognathus mesembrinus named by Peter Wellnhofer in 1987 was assigned to Ornithocheirus by David Unwin in 2003 (making Tropeognathus a junior synonym), but as Anhanguera mesembrinus by Alexander Kellner in 1989, Coloborhynchus mesembrinus by André Veldmeijer in 1998 and Criorhynchus mesembrinus by Michael Fastnacht in 2001.
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In 1881 he designated O. simus the type species of Ornithocheirus and named a new species O. bunzeli.
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Coloborhynchus, "maimed beak", comprised a new species, Coloborhynchus clavirostris, the type species, and two species reassigned from Ornithocheirus: C. sedgwickii and C. cuvieri.
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The type specimen of Ornithocheirus oweni Seeley 1870, CAMSM B 54439, was referred to Lonchodraco(?) microdon in the same study that named Lonchodraco, following a conclusion by David Unwin in 2001, and this species would then be a junior synonym.
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Also unlike related pterosaurs, the teeth of Ornithocheirus were mostly vertical, rather than set at an outward-pointing angle.
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While it has traditionally been classified in the genus Ornithocheirus, it is more likely an azhdarchid, though due to the fragmentary nature of known fossil remains, it is considered a nomen dubium.
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He soon concluded that it was best to create a new genus for the Cambridge Greensand material that he named Ornithocheirus, "bird hand", as he in this period still considered pterosaurs to be the direct ancestors of birds, and assumed the hand of the genus to represent a transitional stage in the evolution towards the bird hand.
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They renamed Pterodactylus sedgwickii Owen 1859 into Camposipterus(?) sedgwickii and Ornithocheirus colorhinus Seeley 1870 into Camposipterus(?) colorhinus.
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Keeping the name Ornithocheirus, he added to it Owen's Criorhynchus, in which however Coloborhynchus was sunk, and to allow for a greater differentiation created two new genera, again based on jaw form: Lonchodectes and Amblydectes.
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Ornithocheirus brachyrhinus Seeley 1870 and Pterodactylus fittoni Owen 1859 share several features with C. cuvieri, and it can therefore not be excluded that they are the same taxon.
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This pterosaur is closely related to Ornithocheirus, and belongs in the family Ornithocheiridae within its own subfamily, Anhanguerinae.
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