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Poekilopleuron
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The most distinctive feature of Poekilopleuron were its forelimbs.
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Using the characters described in this study, Allain defined Megalosauridae as dinosaurs including Poekilopleuron valesdunesis, now known as Dubreuillosaurus, Torvosaurus, Afrovenator, and all descendants of their common ancestor.
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In 2002, when this process had not yet been completed, Ronan Allain, after having dedicated a thesis to it, named the find as a new species of Poekilopleuron: Poekilopleuron? valesdunensis.
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In 2002, Ronan Allain redefined the clade after he discovered a complete megalosaurid skull in northwestern France of species Poekilopleuron.
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The type species of the genus is the original Poekilopleuron valesdunensis, the combinatio nova is Dubreuillosaurus valesdunensis.
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Unlike later Theropoda, whose forelimbs tended toward reduction in length in proportion to the animals' size, Poekilopleuron's were long and, by implication, potent.
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However, the position of Xuanhanosaurus and Poekilopleuron is very unstable, and their exclusion from the analysis gave more resolved and stable cladogram.
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Because the original fossil was destroyed and no other remains of Poekilopleuron have since been found, and also because of its name change, there is much controversy surrounding its classification that cannot be resolved.
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Both Poekilopleuron and Lourinhanosaurus were recovered outside the family, and many taxa within the Metriacanthosauridae were in polytomy.
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The question mark after the generic name already indicated the assignment to Poekilopleuron was tentative.
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Allain also defined two taxa within Megalosauridae: Torvosaurinae was defined as all Megalosauridae more closely related to Torvosaurus than to Poekilopleuron and Afrovenator, and Megalosaurinae was defined as all those that are more closely related to Poekilopleuron.
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By 2005, Allain had come to the conclusion that the new species was not part of Poekilopleuron.
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Unlike Benson et al., 2010, they recovered Poekilopleuron as an afrovenatorine, while Xuanhanosaurus was recovered as the basalmost metriacanthosaurid.
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However, it did not include Dubreuillosaurus (formerly Poekilopleuron valesdunensis), which could affect the results in that region of the cladogram (Carrano et al. 2002). Other more recent and more complete cladistic analyses show Afrovenator in a group of Megalosauridae with Eustreptospondylus and Dubreuillosaurus.
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Allain also defined two taxa within Megalosauridae: Torvosaurinae was defined as all Megalosauridae more closely related to Torvosaurus than to Poekilopleuron and Afrovenator, and Megalosaurinae was defined as all those that are more closely related to Poekilopleuron.
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However, it did not include Dubreuillosaurus (formerly Poekilopleuron valesdunensis), which could affect the results in that region of the cladogram (Carrano et al.
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